1 Hainan Jianfengling Forest Ecosystem National Field Science Observation and Research Station, Research Institute of Tropical Forestry, Chinese Academy of Forestry, Guangzhou 510520, China; 2 College of Life Sciences, Gannan Normal University, Ganzhou 341000, Jiangxi, China; 3 Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China; 4 Departamento de Zoología, Facultad de Biología, Universidad de Santiago de Compostela, 15782 Santiago de Compostela, Spain; 5 Hainan Academy of Forest (Hainan Academy of Mangrove), Haikou 571100, China *Author for correspondence. E‐mail: qye@scbg.ac.cn