1. Bryology Laboratory, School of Life Sciences, East China Normal University, Shanghai 200241, China

2. Shanghai Institute of Eco‐Chongming (SIEC), Shanghai 200062, China

3. Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural Sciences, Shenzhen 518124, China

4. State Key Laboratory of Plant Diversity and Specialty Crops, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China

^*Author for correspondence. E‐mail: fangyuhan@scbg.ac.cn