1State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China 2University of Chinese Academy of Sciences, Beijing 100049, China 3Shenzhen Key Laboratory for Orchid Conservation and Utilization, The National Orchid Conservation Center of China and The Orchid
Conservation and Research Center of Shenzhen, Shenzhen 518114, China 4Key Laboratory of National Forestry and Grassland Administration for Orchid Conservation and Utilization at the College of Landscape
Architecture, Fujian Agriculture and Forestry University, Fuzhou 350002, China 5College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510640, China 6Fujian Colleges and Universities and Engineering Research Institute of Conservation and Utilization of Natural Bioresources, College of
Forestry, Fujian Agriculture and Forestry University, Fuzhou 350002, China 7Sino‐Africa Joint Research Center, Chinese Academy of Sciences, Wuhan 430074, China