1 Institute of Entomology, College of Life Sciences, Nankai University, Tianjin 300071, China; 2 Co-Innovation Center for the Sustainable Forestry in Southern China, College of Forestry, Nanjing Forestry University, Nanjing 210037, China †These authors contributed equally to this paper. *Authors for correspondence. Huaijun Xue. E-mail:xuehj@nankai.edu.cn;Wenjun Bu. E-mail:wenjunbu@nankai.edu.cn