1. Department of Environmental Systems Science, ETH Zurich, Schmelzbergstrasse 9, Zurich CH-8092, Switzerland 2. College of Life Sciences, Beijing Normal University, Beijing 100875, China 3. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China Author for correspondence. E-mail:pengjuan.zu@gmail.com
