An attempt is made to elucide some pronounced changes in the classification ofLiliaceae sensu lato in the last 60 years. The changes in the classification of Liliaceae s. l. have to involve the classification of its related families. In order to understand the different treatments of it by Engler ＆ Prantl (1930), Hutchinson (1934), Dahlgren et al. (1985, embodies G. Dahlgren, 1989), Takhtajan (1987), and Thorne (1992), a concise table of approximate concordance of taxa (with tribe as a main basic unit) of the order Liliiflorae sensu Engler ＆ Prantl with those recogizied in the other four systems is given. The considerations and the taxonomic characters adopted by Hutchinson, who may be the first author to change drastically the classification of Liliaceae sensu lato, are more detailedly explained than those in Dahlgren et al. , Takhtajan, and Thorne's systems. Comments on the benefits of narrower and broader circumscription at family level are made. The present author does not agree with elevating all subfamilies or satellite groups to the rank of family if the segregated families remain the closest relatives without the change in systematic position. Different hypotheses of the origin of the monocotyledons proposed by Hutchinson, Takhtajan and Dahlgren et al. are also introduced. Although the opinion of Dahlgren et al. (1985) has not been accepted by many authors, their peculiar methodology is worth noting. On the basis of the distribution of character states common to the monoctyledons and the dicotyledons they postulate a hypothetical taxon. Then the derivation of an ancestral monocotyledon from this hypothetical taxon might only have involved a few steps. Such an ancestral monocotyledon does not exist, but the closest extant plant is perhaps Trichopus (Trichopodaceae, Dioscoreales). Certainly Tichopus is not a direct link between the monocotyledons and the dicotyledons, but it exhibits a number of features that are also found in Annonales (superodeer Magnolianae). Thus the Dioscoreales are favoured by them as the least derived monocotyledons. They place the Dioscoreales at the beginning of their system as the first order of the superorder Lilianae. The author gives some suggestions for investigations of Liliaceae s. l. in China today. Among them the extensive studies (in morphological, chromosomal and molecular aspects) of some selected groups, such as Lilium, Monocharis, Paris etc. , seems more realistic because they are distributed in N. Hemisphere. It is easy to collect the materials. Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
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The geographical distribution of Liliaceae (s. str. ) is analysed on the basis of the floristic regions proposed by Takhtajan. Some conclusions may be proposed as follows: 1.Liliaceae (s. str. ) consists of nine genera and about 513 species, distributed primarily in the north temperate zone. Statistics shows clearly that the Irano-Turanian Region is most abundant in number of species, The Eastern Asian Region with total nine genera of the family is the diversity centre of Liliaceae (s. str. ). 2. The distribution patterns of the genera may be divided into: 1 ) North temperate distribution pattern: Lloydia, Erythronium, Fritillaria and Lilium; 2) Old world temperate distribution pattern: Gagea and Tulipa; 3) West Asia to Himalayas and Southwest China distribution pattern: Notholirion; 4) East Asia distribution pattern: Cardiocrinum and Nomocharis. 3. The Sino-Himalayas is one of the key regions in studying the evolution of Liliaceae (s. str. ) All nine genera occur in the Eastern Asian Region with most species distributed in Southwest and Northwest China. Chorologically, five genera (Fritillaria, Lilium, Cardiocrinum, Nomocharis and Notholirion) of the Lilieae are overlapped each other in the Sino-Himalayas, showing its diversity centre in this region. The Lilieae is a main stock in the Liliaceae (s. str. ), The genus Nomocharis in this tribe may have been newly differentiated from Lilium in the course of the uplift of Qinghai-Xizang Plateau, a view also supported by Xie Xiao-yang et al.. The place of its origin was considered to be in the southern part of the Hengduan Mountains. 4. The distributions of some species in Liliaceae (s. str. ) seem to be significant for dividing some floristic regions: 1 ) Some species of Fritillaria (F. unibracteata Hsiao et K. C. Hsia, F. przewalski Maxim. ex Batal. , F. crassicaulis S. C. Chen, F. cirrhosa D. Don. , F. delavayi Franch. , F. dajinensis S. C. Chen, F. davidii Franch. , F. sinica S. C. Chen and F. sichuanica S. C.Chen) are only distributed in Sino-Himalayan forest subkingdom, while others (F. taiparensis P. Y. Li, F. yuzhongensis S. C. Chen, F. monantha Migo, F. anhuiensis S. C. Chen et S. F. Yin, F. thunbergii Miq. , F. maximowizii Freyn and F. ussuriensis Maxim. ) are restricted to Sino-Japan forest subkingdom. They show a clearly demarcation line between the two subkingdoms, which is identical with what proposed by Wu Cheng-yih. 2) The Eastern Asian Region can be divided into two subkingdoms on the basis of the distribution pattern of the genus Cardiocrinum; C. giganteum (Wall.) Makino and C. gigateum var. yunnanense Leichtlin ex Elwes are restricted to Sino-Himalayan forest subkingdom. C. cathayanum (Wilson) Stearn and C. cordatum (Thunb.) Makino are only found in Sino-Japan forest subkingdom. 3) The distributions of Gagea pauciflora Turcz. , G. triflora(Ledeb.) Roem. et Schult. G. hiensis Pasch, Lloydia tibetica Baker ex Oliver, L. oxycarpa Franch. and L. flavonutans Hara are indicative of a demarcation line between Irano-Turanica Region and Eastern Asian Region. 5. The genus Notholirion occurs in the Eastern Asian Region and Irano-Turanian Region, showing the relationships between the two regions and also between the Chinese flora and Ancient Mediterraneam flora.

Pollen morphology of three species in the genus Maharanga and 12 species in the genus Onosma , belonging to the family Boraginaceae, was examined by means of light microscope and scanning electron microscope. Pollen grains of the genus Maharanga are dumbbell-shaped or cocoon-shaped, medium-sized, P/E 1.6--1.67 ratio, tricoporate with lalongate ora,and spinulate,while those of the genus Onosma are subprolate or ovoid, P/E 1--1.23 ratio, tricolporate or trisyncolporate with lolongate ora, rugulate-microgranulate or tuberculate. The pollen morphology supports the separation of the genus Maharanga from the genus Onosma (sensu lato). Noteworthily, the three colpi of syncolporate pollen in some members of the genus Onosma converge in one pole or even two poles.

The present paper deals with the morphology, classification, systematic arrangement and geographical distribution of the genus Ribes L. from China. Many botanists made some investigations on the systematic position of this genus, but their views have been controversial for a long time. It was recognized as a genus of Saxifragaceae by some taxonomists, while others treated it as an independent family, i. e. Grossulariaceae or Ribesiaceae, based on external morphological and pollen characteristics. The present paper supports the treatmemt of the genus Ribes L. as a member of the Saxifragaceae. Through detailed analysis of the characters of main organs, the evolutionary trends are suggested: the flower has evolved from large and bisexual to small and unisexual; the calyx from tubular or cylindric to campanulate, cupuliform, pelviform to discoidal, even to rotate; the inforescence from a raceme to a corymb or a umbell, then reduced to several flowers or a single flower;the bract from small to large and its shape from ovate, nearly rounded to elliptic, oblong,lanceolate or linear. Based on the data from other authors, the evolutionary tendencies of the pollen morphology are as follows: the aperture type from zonocolporate to pantoporate; the shape of ectoaperture from colpate to porous and the number of endoapertures per ectoaperture reduced from 2 to 1. The analysis of morphological and pollen characteristics shows that the subdivision of Ribes into 4 subgenera in Rehder's system is reasonable and is accepted by the present author, with some important revisions in the subdivisions of subgenera and sections, and systematic arrangements. In addition, the systematic arrangement of the subgenera and sections presented here in this paper is rather different from that in Rehder's system from the phylogenetic point of view. Fifty-nine species and 30 varieties are recognized from China, belonging to four subgenera, 10 sections and 15 series. Five series are described as new. Ribes L. is basically a genus of North Temperate area of the four subgenera, 14 sections and more than 160 species in the world, four subgenera, 10 sections and more than 80 species are recorded in Asia. Among them, four subgenera, 10 sections and 44 species, including primitive, intermediate and advanced taxa of phylogenetic importance are concentrated in Eastern Asiatic region. It is clearly shown that the Eastern Asiatic region is not only the frequency center, but also the diversity center of the genus Ribes L. at the presenttime. In this floristic region the greatest number of species and endemics exist in China, i. e. in Sichuan Province and the adjacent regions, including W. Sichuan, NW. Yunnan, SE. Gansu, S. Shaanxi, W. Hubei and W. Hunan, Guizhou, E. Xizang and E. Qinghai, where the genus also reaches its highest morphological diversity. According to the above fact, we tend to infer that the Hengduan Mountains region in the Sino-Himalayan Forest Subkingdom and the west part of the Sino-Japan Forest Subkingdom is the most important region for the occurence and differentiation of the genus Ribes L. Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug.
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